Will PCIe x1 x4 cards work in x16 slot?

pcie slot x16
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On current Intel single socket systems a PCIe switch is required to get 4 full X16 slots for GPU's The alternative possibility for 4 GPU's on

pcie expansion slot slot only supports PCIe x16 which comes from the Z590 chip Is there a problem with the first PCIe slot on this new motherboard?

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